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Emotional Neural Systems

From Aristotle’s earliest attempts to list emotions and passions up to the modern taxonomies, the grouping criteria have been confused and disparate, even though, rational psychology had already postulated the distinction between primary and secondary emotions, or between basic and compound, for it had been observed that many of the emotional states maintained certain similarities amongst themselves. It was a blind proceeding, without physiological bases, something that led in some cases, like that of Spinoza, to confound pleasure and pain with basic emotions. In the works of Paul Broca (1878), James Papez (1937) and Paul Mclean (1952), it had been proposed that human emotions were linked to specific brain parts, the topo-biological set that we have called limbic system, or visceral brain.[1] From this line of research, it was developed the so called categorical approach to emotions, which proposed the existence of a small group of emotions from which the others were derived. Such a proposal was at the same time based on the traditional philosophical approach and on Broca, Papez and Mclean’s theses of brain partition, as well as on the analysis of the emotional expressions of behavior. The categorical theses assume that certain affective processes, such as fear, rage, joy or sadness, emerge from stable intrinsic brain systems that can be clarified at a biological level.[2] The approach that I will be following hereby is categorical, for the existence of different intrinsic neural systems is a corollary of the process of categorization which leads to the primary consciousness in the model of Neural Darwinism. By this, I do not mean to say that other conditionings apart from the evolutionary and epigenetic may not intervene in human emotions, although it is necessary to clarify first the psycho-biological dimension, mainly when social behavior itself, which introduces new conditionings in the basic emotions, seems to have its origin in the emotional protocols. However, I do consider that socio-constructivist approaches, which pretend to understand emotions by means of processes of mere linguistic analysis, cannot but return to the explicitation of the old mythologies in which local ethics are established.
            Obviously, from onto-theological standpoints, the hypothesis of a continuity between our emotions and those of all other mammals is merely rejected on metaphysical grounds, and the profound similarities and clear continuities observed at a neurochemical level –when not identical phenomena- are discarded based on ontological discontinuities produced by the action of supernatural phenomena.[3] An objection could be made, from this point of view, saying that the approach to emotions that I follow here confounds instinct with emotion, that human emotions are more complex phenomena than a mere sensation of fear. Mythological systems, and the arts based on them, are sufficient proof of the difference in complexity between animal and human emotions, however, to proclaim the autonomy of our emotions with respect to the natural world is nothing but a variation of the traditional theological theses about the origin of man. Here I accept the proposal of William James about the imperceptible overlapping that occurs between instinctive reactions and emotional expressions.[4] The distinction between instinct and emotion is usually established based on two criteria: the difference between animal and human, and the difference between mechanical or unconscious processes, and volitional or conscious processes. The first distinction, maintained nowadays by authors of social constructivism,[5] or from different religious positions, is based on the idea that humans do not have instincts, and that affections are acquired in learning processes. Such proposals are a variant of the Cartesian theories which presented animals as mere automata, while making man a divine being in his origin and his functioning. Genetic science, the theory of evolution and ethology refute a radical distinction and confirm the Jamesian one. However, this attitude of rejection cannot come as a surprise. After all, what we identify as constituent of our personality are some specific emotional tendencies and a given emotional and sentimental history, so the fact that our psychological intimacy and our most profound values could have their origin in a group of physiological conditionings, and not in a plan that was outlined from eternity for my individuation, can be taken as metaphysically and morally offensive. The second distinction that separates instinct from emotion based on the consciousness of the process, is invalidated by the difficulties that the very same concept of consciousness has. Primary consciousness is a property of the neurological processes of categorization in mammals, and is not clear that a difference could be established at this level between experimenting an instinct and knowing that one is experimenting that instinct (which it would be, according to this proposal, the emotion). Human consciousness is not only primary, but it incorporates symbolic linguistic processes, and in this respect, emotions themselves are only neural processes concurrent with others that we would not consider as such. The definition of emotion based on the theses of Neural Darwinism is not separable from the definition of primary consciousness, but such a concept of emotion would be in many cases indistinguishable from that of instinct in accordance with the ambiguous categorizations inherited from philosophy and psychology.
            The link of a small group of emotions to specific neuroanatomical parts is sustained, in fact, on rather crude experimental bases. Thus, a temporal lobotomy performed in cats, apes or people, produces sexual hyperactivity, hyper-orality and a decrease of fear, whereas a lesion in the frontal lobe makes animals more placid, or a septal lesion makes them aggressive, and these emotional states are repeated one experiment after the other.[6] Obviously, when we study the brain looking for a topo-biology of emotions, we already start with a preliminary idea of emotion, and during experimentation we find more or less diffuse concepts which we had already carried into the experiment. It is also true that the specific emotions that we have encountered, expressed in lesions or lobotomies, have always coincided with emotional processes which since Aristotle were considered basic passions, such as rage or fear.
            There is no widespread agreement about basic emotions in categorical theories. Here, I will adopt the model proposed by Panksepp, organized around strictly naturalist postulates and fundamentally derived from neuroscientific studies in animals. Panksepp began with four categories of animal behavior, fear, rage, appetitive desire and sociability, with the goal of finding their topo-biology, but he found during the experimental process a total of seven topologies or emotional neural systems, with their corresponding neurochemical circuits, which expanded the starting categories. These seven topologies are brain structures whose actions are linked to behaviors that we identify with vital motivation, affective aggression, fear, sexual motivation, socio-organic behavior, maternal care, and ludic behavior.[7] These systems allow isolated stimuli, they function physiologically in an independent manner at a neurochemical level in experimental conditions, even though, by coexisting within the neurological system of an organism that interacts with its environment, they overlap and influence one another in different ways, originating the complexity of the emotional world. The study of these systems in mammals allows us to understand them in their primitive and derived (or mixed) forms, with the integrative mutual causations of limitation and potentiation. I will call primary to the primitive emotions of the group of the seven aforementioned systems, and n-ary to the mixed emotions.
     On a preliminary basis we can define emotions -following Panksepp’s criteria- as psycho-neural processes that have special influence on the control of vigor and the formation of patterns in actions that are performed amongst animals, or between animals and objects, during circumstances which are especially important for survival.[8] If we define intelligence in evolutionary terms as the ability of a living being to respond with flexibility and successfully to new vital scenarios, the emotional functioning of the brain is already a form of intelligence.[9] Neuroscience speaks about two types of neural activity which are considered as intelligence in this evolutionary sense: the cognitive and the socio-emotional. The executive functions of cognition imply the maintenance and manipulation of information which is indispensable in order to deal with situations in which the appropriate response is not dictated by the actual information of the stimulus. It is customary to include as actions of the cognitive neural systems the working memory, the attention, the representation and planning of objectives, the monitoring of responses and error detection.[10] Whereas cognitive intelligence is associated with reasoning, the emotional and the social are linked to perception and immediate processing. Cognitive intelligence is strategic and cortical, whereas the emotional is tactical and limbic, useful for immediate behavior, and better prepared for survival and adaptation, even though, when we speak of a human being these argumentations no longer make much sense, for the adaptations that have allowed his domination over the entire animal realm have originated from a development without parallel of his cognitive capabilities. A different matter is that at moments of vital stress the emotional influences of the subcortical system can be stronger than the control exerted from the most evolved parts of the brain, as we see and experience in wars and liminal situations, in which the basic emotions impose the rationality of survival. The link of emotions to the homeostasis of the organism turns them into protocols of quick response to liminal environmental stimuli, a speed which is irrelevant in strategic and cognitive decision making, which involve slower processes in which a higher quantity of information is contrasted. Nonetheless, emotions are not limited to be momentary responses, for the neural activity of the emotive systems lasts more than the precipitous circumstances, and the distinction between strategic and tactical decision loses its meaning.  



[1] Consisting on the limbic lobe, the dental gyrus, the amygdala, the septal nuclei, the mammilar bodies, the anterior thalamic nucleus, the olfactive bulbs, and myelinated axon bundles. New experimental data from 2004 (See Dalgeish, T. The Emotional Brain. Nature Reviews Neuroscience. Web.) include as part of the emotional system also the prefrontal cortex, the ventral striatum, the insula, the anterior cingulate and the cerebellum.
[2] Cf. Panksepp. Op. Cit. p.44.
[3] Like the declaration that Pope John Paul II made at the Pontifical Academy of Sciences on the 22nd of October of 1996, proclaiming a discontinuity between human and ape from a certain historical moment in which the divine intervention breathed human soul into an animal linage. 
[4] To James, all which stimulates an instinct also stimulates an emotion. Cf. William James. The Principles of Psychology. Vol.II. Ed. Cit. p.442.
[5] See the theses of G. Mandler. Mind and Body: The Psychology of Emotion and Stress. Norton. New York. 1984.
[6] Cf. Panksepp. Op. Cit. p.79.
[7] Panksepp. Ibid. p.p.52-54. The emotion which I have called socio-organic or social is the one that Panksepp calls panic, a poorly fortunate name due to its etymology (a type of fear related to Pan, the Greek god of nature), even though it denotes the same concept in affective neuroscience.
[8] Cf. Panksepp. Ibid. p.48.
[9] The definitions of traditional psychology, which since Wundt made emphasis on intelligence as a processes of logical thinking (Wundt. Elements de Psychologie Physiologique. Félix Alcan. Paris. 1886. Tome Second. p.365.)
[10] See Gazzaniga, Michael S., Ivry, Richard B., Mangun, George R. Cognitive Neuroscience. The Biology of the Mind. Norton. New York. 2009.

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